,+&ơ11*
751*,+&63+0
KHOA SINH 0é,751*

3+$17++8<175$1*

1*+,ầ1&8
72
&+3+0
6,1++&
1*'1*
;/ộ0ễ1&$75ơ0
&+7751*106ẹ7ậ0

j1QJ
1P


,+&ơ11
G
751*
,+&63+0
KHOA SINH 0é,751*

3+$17++8<175$1*

1*+,ầ1&8
72
&+3+0
6,1++&
1*'1*

;/ộ0ễ1&$75ơ0
&+7751*106ẹ7ậ0

NgjQK
&{QJQJKVLQKKF

&ẩ1%+1*'1
7K61*8<17+%ậ&++1
G

j1QJ
1P
2018


/Ӡ,&$0Ĉ2$1

7{L[LQFDPÿRDQÿk\OjF{QJWUuQ
&iFV͙OL͏XN͇WTX̫WURQJOX̵QY
DLF{QJE͙WURQJḘWNǤF{QJWUuQKN

TiFJLҧOXұQYăQ

3+$17+ӎ+8<ӄ175


/ͤ,&̪0ˮ1

Ĉ͋KRjQWKjQKÿ˱ͫFNKyDOX̵QQj\
F{JLi

o K˱ͣQJG̳Q7K61JX\͍Q7K͓%tFK+
K˱ͣQ
JG̳QYjWUX\͉Qÿ̩WNL͇QWKͱFNL
WUuQKWK͹FKL͏QNKyDOX̵Q
(P[LQFKkQWKjQKF̫P˯Qÿ͇QFiF
±0{LWU˱ͥQJ
±
Ĉ̩LK͕F6˱3K̩P
±Ĉ̩LK͕FĈj1̽QJÿmW̵QWuQK
NL͇QWK͹FFKRHPWURQ
JTXiWUuQKK͕FW̵SW̩LWU˱
&X͙LFQJHP[LQF̫P˯QJLDÿuQK
WKͥLJLDQHPOjPNKyDOX̵Q
;LQFKkQWKjQKF̫P˯Q


0&/&

08
.....................................................................................................................1
7ậ1+&37+,7&$7ơ,
..........................................................................1
0&7,ầ8&$7ơ,
.....................................................................................3
ộ1*+$&$7ơ,
.......................................................................................3
ộQJKDNKRDKF
............................................................................................3
&+1*,71*48$17ơ,/,8
.....................................................................4

1.1. T1*48$199
I SINH 97
.......................................................................4
7QJTXDQYYLNKXQ
Bacillus ....................................................................4
7QJTXDQY[DNKXQ
Streptomyces ............................................................5
7QJTXDQYQPPFWUQJ
Phanerochaete chrysosporium....................6
71*48$191*ơ1+109ơ48<75ẻ1
758<17+1*
......................................................................................................7
7QJTXDQYQJjQKQPQFW
..............................................................7
7uQKKuQKVQ[XWYjWLrXWK
..........................................7
7uQKKuQKVQ[XWYjWLrXWK
........................................8
6OFYJLiWUFWtQKVLQK
...9
*LiWUGLQKGQJFDQPVz
.....................................................................9
*LiWUNLQKWFDQPVz
............................................................................9

FWtQKVLQKKFFDQPVz
......................................................................10
7QJTXDQYFKSKPVLQKKF
.......................................................................13
.KiLQLPFKSKPVLQKKF

.....................................................................13
&VNKRDKFFDYLFVGQJ
KF
.........................................................................................................................13
7uQKKuQKQJKLrQFXQJGQJ
[OờFFK
WUQJQP
..............................................................................................................14
&+1*,71*1,'81*9ơ3+1*
..17
,71*1*+,ầ1&8
...........................................................................17
LWQJ
....................................................................................................17
DLPYjWKLJLDQQJKLrQF
..............................................................17
2.1.3. NguyrQOLXKyDFKWYjWKLWE
...............................................................17
1,'81*1*+,ầ1&8
..............................................................................18
3+1*3+ẩ31*+,ầ1&ề8
......................................................................18
3KQJSKiSSKkQOSYjJLJL
............................................................18
3KQJSKiS[iFQKNKQQJS
..................................19
3KQJSKiSQKOQJYLVLQK
...........................................................19
3KQJSKiSVQ[XWFKSKPY
.....................................................20

3KQJSKiSPQFD
.............................................................................21
3KQJSKiSiQKJLiFPTXDQ
....................................................22
3KQJSKiSiQKJLiQQJVXW
......................................22
3KQJSKiSVOờVOLX
...........................................................................22
&+1*.7489ơ%,1/81
..............................................................23
.7483+ặ
1/3&ẩ&&+1*969
....................................................23
3KkQOSYLNKXQ
Bacillus: .........................................................................23
3KkQOS[NKXQ
Streptomyces .................................................................24
3KkQOSQPPFWUQJ
..............................................................................25


.ӂ748Ҧ;È&Ĉӎ1+.+Ҧ1Ă1*3+Ỉ1*,
.................27
3..KҧQăQJSKkQJLҧLFHOOXORVHFӫD
Bacillus .................27
.KҧQăQJSKkQJLҧLFHOOXORVHFӫD
Streptomyces ...............28
.KҧQăQJSKkQJLҧLFHOOXORVHFӫ
.....................29
.ӂ748Ҧ;È&Ĉӎ1+7Ë1+ĈӔ,.+È1**

1Ҩ06Đ7Ë067

..................................................................................................30
.7481*+,ầ1&861;87&+3+
..................31
.7487&/$17&$106$8
%1*&+3+0
.................................................................................................32
.+26ẩ77/1+,0&$&ẩ&%&+
.......................................33
11*68710%ơ21*7ậ0
...............................................................33
&+1*.
7/819ơ.,11*+
............................................................35
.WOXQ
..............................................................................................................35
.LQQJK
............................................................................................................35
7ơ,/
,87+$0.+2
.........................................................................................36


'$1+0Ө&&+Ӳ9,ӂ77Ҳ7

&)8&RORQ\)RUPLQJ8QLW
ÿѫQYӏ
9699LVLQKYұW
&36+&KӃSKҭPVLQKKӑF

&3&KӃSKҭP
Ĉ&ĈӕLFKӭQJ
676z7tP
&7&{QJWKӭF


'$1+0Ө&%Ҧ1*%,ӆ8
STT
Bҧ
ng 3.1

7rQE
ҧng
Trang
+uQKWKiLNKX̱QO̩FFͯDFiFFK
24
SKkQO̵SWͳP̳Xÿ̭W

Bҧ
ng 3.2

+uQKWKiLNKX̱QO̩FFͯDFiFFK
25
SKkQO̵SWͳP̳Xÿ̭W

Bҧ
ng 3.3

+uQKWKiLNKX̱QO̩FFͯDFiFFK
26

ÿ˱ͫFSKkQO̵SW
ͳP̳Xÿ̭W
9zQJSKkQJL̫LFͯDFiFFKͯQJ%DF
27

Bҧ
ng 3.4

cellulose

Bҧ
ng 3.5
Bҧ
ng 3.6

Bҧ
ng 3.7
Bҧ
ng 3.8

9zQJSKkQJL̫LFͯDFiFFKͯQJ6
28
WtQKFHOOXORVH
9zQJSKkQ
JL̫LFͯDFiFFKͯQJQ̭PPͭ
Fy 29
KR̩WWtQKFHOOXODVH

K͇WTX̫ÿ͙LNKiQJJLͷDQ̭PPͭF
30

ttP
6RViQKV͙O˱ͫQJEjRW͵WKXÿ˱ͫF
31
PHQ[͙SW̩RFK͇SḴP

Bҧ
ng 3.9
Bҧ
ng 3.10
Bҧ
ng 3.11

7͙Fÿ͡ODQW˯VDX
QJj\
32
T͑O͏QKL͍PWUrQFiFE͓FK
33
1ăQJVX̭W
VLQKK͕
FFͯDQ̭PEjRQJ˱WtPW
33
QJj\


'$1+0Ө&+Ỵ1+Ҧ1+

STT
7rQE
ҧng
Trang

+uQK
+uQK̫QKQKX͡P*UDPP͡WV͙FK
4
+uQK 6˯ÿ͛WyPW̷WTX\WUuQKWU͛
11
+uQK 6˯ÿ͛WyPW̷WTX\WUuQKW̩R
21
+uQK
3.1
+uQKWKiLNKX̱Q
O̩FFͯDFiFFKͯQJYLNK
24
SKkQO̵SWͳP̳Xÿ̭W
+uQK
3.2

+uQKWKiLNKX̱QO̩FFͯDFiFFK
25
SKkQO̵SWͳP̳Xÿ̭W

+uQK
3.3
+uQKWKiLNKX̱QO̩FFͯDFiFFK
26
ÿ˱ͫFSKkQO̵SWͳP̳Xÿ̭W
+uQK
3.4
9zQJSKkQJL̫LFͯD
FiFFKͯQJ%DFLOOXVFy
27

cellulose
+uQK
3.5
9zQJSKkQJL̫LFͯDFiFFKͯQJ6
29
WtQKFHOOXORVH
+uQK
3.6
9zQJSKk
QJL̫LFͯDFiFFKͯQJQ̭PP
Fy 30
KR̩WWtQKFHOOXODVH

+uQK
3.7
K͇WTX̫ÿ͙LNKiQJJLͷDQ̭PPͭF
Vz 30
ttP


08

1. 7ậ1+&37+,7&$7ơ,
1POjWKFSKPJLjXFKWGLQKG
YLWDPLQQKYLWDPLQ%%33ôYj
FQJQJPFKYjNK{QJOjPWQJO
1PFzQF
QKLXWiFGQJGFOờNKiSKRQJSK
FDFWKNKiQJXQJWKYjNKiQJY
PFKKQJPiXFKQJSKyQJ[

EjRJDQDQWKQUW
FyOLFKRYLFLXFKQK
WUXQJQJ
7UrQWKJLLFyNKRQJORjL
Q QJRQ Yj F QX{L WUQJ QKkQ WR
FjQJSKiWWULQFiFORLQPFW
QOjPFQKQ
PQPEjRQJQPOLQKFKLFiFOR
1FWDFyWLPQQJOQYVQ[
QJXQQJX\rQOLXWUQJQPSKRQJSK
LX NLQ WKӡL WLӃW
QKLӅX
WKXұQ
FKӫQJ
OӧLORҥL
FKR S
WUӗQJ QҩP TXDQK QăP &K~QJ WD ÿm Fѫ
JLӕQJYjVҧQ[XҩWQҩPÿӕLYӟLFiFO
QJj\FjQJUӝQJPӣ&KtQKYuYұ\QJj
baQ KjQ
K 4X\ӃW ÿӏQK
- TTg, ÿѭD
4Ĉ
QҩP ăQ QҩP GѭӧF
PөFVҧQSKҭPTXӕFJLDÿѭӧFѭXWLrQÿ
7ӱ&KtQK3Kӫ1ѭӟF&ӝQJ+zD;m+ӝL&K
1ѭӟFWDVҧQ[XҩWNKRҧQJORҥLQ
\ӃXWU
ӗQJ
QҩPUѫPQҩPPӝFQKƭ&

iFWӍQKSKtD%ҳFFKӫ\ӃX
QҩPOLQKFKLYӟL6ҧQOѭӧQJQҩPNKRҧQ
-  WULӋX 86' NK{QJ WtQK [XҩW NKҭX
WҩQQҩPUѫPWҩQ
QҩPVzWҩQQҩP
OLQKFKLWҩQFiFORҥLQҩPNKiF
FKkPQҩPQJӑFFKkPNKRҧQJWҩQ
0ӝWWURQJQKӳQJORjLQҩPSKәELӃQ
1DPÿyOjQҩ
OjPӝWORjLQҩPăQÿѭӧFFyJLiWU
ӏGLQKGѭӥQJFDRFyY
JLiWUӏGLQKGѭӥQJFDRFXQJFҩSFi
FKӭDQKLӅXSURWHLQJOX[LWYLWDPLQ
GӉKҩSWKөEӣLFѫWKӇFRQQJѭӡL+j
QJQҩPVzWӯ
1ҩPVzKRjQWRjQFyWKӇWKD\W
WӯÿӝQJYұW'RÿyQ
ҩPVzFzQÿѭӧFJӑLOj³UDX
´³WKӏWVҥFK´
VӱGөQJQKѭQJXӗQFXQJFҩSSURWHLQF
KӑFFiFFK
ҩWGLQKGѭӥQJYjYLFKҩWFyOӧL
1


ÿѭӧFFKX\ӇQKyDWKjQKQăQJOѭӧQJFKR
NLrQJ´GjQKFKRFiFEӋQKQKkQWLӇX
FyWKyLTXHQăQFKD\ĈӕLYӟLQJѭӡL
WKӇFiFPyQăQ
QҩP JL~S SKөF KӗL VLQK OӵF QKDQK FK

NK{QJ ÿzL KӓL FҫX Nu Pj YүQ UҩW QJR
OXӝF«YӯDFyWiFGөQJEәGѭӥQJYӯD
QҩP Vz
ѭӧF
Fy
[HP
ÿ
Oj WKjQK SKҫQ TXDQ WU
FKX\ӇQ QăQJ OѭӧQJ Yj FiF WKjQK SKҫQ
1JRjLUDFK~QJFyYDLWU
zFKtQKWURQJKӋPLӉQGӏFK
WLQKYjVLQK
WULӇQ
%rQFҥQKÿyNKRҧQJPӝWQӱDKӧSFKҩ
FDUERQWURQJVLQK
WUrQPһWÿҩWOjFHOOXORVHFKLӃPWӟ
±NKӕLOѭӧQJNK{
YұW 7ҩW Fҧ VҧQ SKҭP VLQK NKӕL VӁ ÿ
ÿѭӧFFXQJFҩSEӣLYLVLQKYұW+ӋW
FKұ
PYjNK{QJKRjQWRjQ7X\QKLrQW
YLVLQKYұWWURQJGҥFӓEzFyWKӇS
±FHOOXORVH9
YLVLQKYұWSKkQJLҧLFHOOXORVHÿmY
Yjӣ9LӋW1DPQJK
LrQFӭXӢ
ҨQĈӝ%HKHUDYjFV

ÿmSKkQ
YL NKXҭQ SKkQ JLҧL FHOOXORVH Wӯ ÿҩW

Micrococcus spp., Baccilus VSSYj
Pseudomonas VSSӣ7UXQJ4
/LQJ /LDQJ Yj FV 
 ÿm SKkQ
OұS O
FHOOXORVHӢ9LӋW1DP+j7KDQK7RjQ
&DR1JӑFĈLӋS
/r3KҥP7ѭӡQJ
VLQKYұWSKkQJLҧLFHOOXORVH
0һWNKiF
9LӋW1DPOjPӝWQѭӟF
JLjXWLӅPQăQJYӅOkPQJKLӋSGRÿy
QJXӗQSKӃWKҧLQ{QJ
±OkPQJKLӋS
EmPtDUѫPUҥPҥWFѭDUҩWGӗLGjR
YLӋFWUӗQJQҩP+LӋQQD\YLӋF[ӱOê
WK{QJ WKѭӡQJ QJѭӡL GkQ YүQ ӫDR
PҥW
Yj
Fѭ
WKӡLJLDQӫNpRGjL7ӯQKӳQJYҩQÿӅ
KLӋQÿӅWjL
³1JKLrQFӭXWҥRFKӃSKҭPVLQKK
[ӱOêPQF
WUjPOjPFѫFKҩWWUӗQJQҩPVzWtP´

2


2. 0&7,ầ8&$7ơ,

7X\Q FKQ F
YL FiF
NKXQ
BacillusFKQJ
VS [ NK
Streptomyces VSQPPFWUQJ
Phanerochaete chrysosporium&yKRW
SKkQJLLFHOOXORVHFDRFSKkQO
YjRVQ[XWFKSKPYLVLQKQKP[
WtP
3. ộ1*+$&$7ơ,
3.1. éQJKƭDNKRDKӑF
.ӃW TXҧ FӫD
SKҫQ
ÿӅ WX\ӇQ
WjL JyS
FKӑQ
YL
ÿѭӧF
NKXҭQ
F
Bacillus VS [ҥ
Streptomyces
NKXҭQ
VS QҩPFy
PөF
NKҧ
WUҳQJ
QăQJ
JLҧLFHOOXORVHPҥQK

WӯÿyWҥRFKӃSKҭPYLVLQK
PQFѭDWUjP
WUѭӟFNK
i
WUӗQJQҩPVzOjYҩQÿӅFҩSWKLӃWYj
3.2. éQJKƭDWKӵFWLӉQ
6ҧQSKҭPFӫDÿӅWjLFyWKӇӭQJGө
QkQJ FDR QăQJ VXҩWYj FKҩW OѭӧQJ WU
JLҧP QJKqR
WKLӋQ
YjÿӡL
FҧL VӕQJ QKkQ GkQ ĈӅ
KLӋQQD\OjӭQJGөQJF{QJQJKӋVLQK

3


&+ѬѪ1*,7Ә1*48$17¬,/

1.1. 7Ә1*48$19ӄ9
I SINH VҰ7
1.1.1. 7әQJTXDQYӅYLNKXҭQ
Bacillus
9LNKXҭQ
Bacillus OjQKӳQJYLNKXҭQ
*UDPGѭѫQJWKXӝFF
QӝL EjR Wӱ KuQK RYDQ Fy NKX\QK
Bacillus ÿѭӧF
KѭӟQ
SKkQELӋWYӟLFiFORjLYLNKXҭQVLQ

KuQKTXHVLQKWUѭӣQJGѭӟLÿLӅXNLӋQ
7Ӄ
EjR
Bacillus FyWKӇÿѫQKRһFFKXӛLYjFKX\Ӈ
QăQJVLQKEjRWӱQrQYLNKXҭQ
Bacillus FyWKӇWӗQWҥLWURQ
GѭӟL FiF ÿLӅX NLӋQ NKiF QKDX Yj UҩW
ÿѭӧF SKkQ OұS
JXӗQ
WӯNKiF
UҩW QKLӅX
QKDX QKѭ
Q
ÿҩW
WKӭFăQVӳDQKѭQJFKӫ\ӃXOjWӯ
NǤ&Yj1>@
7ҩW Fҧ FiFORjLWKXӝF
Bacillus ÿӅX FyNKҧ
FKL
QăQJGӏ
KRҥL VLQK QKӡ NKҧ QăQJ
KӧS FKҩW
SKkQKӳX
JLҧL
Fѫ
QK
ÿѭӡQJDFLGDPLQDFLGKӳXFѫ0ӝ
WҥR WKjQK JO\FHURO Yj
Bacillus
EXWDQHGLRO

megaterium WKu P
NK{QJFҫQFKҩWKӳXFѫ ÿӇVLQKWUѭӣQ
vitDPLQ%+ҫXKӃWÿӅXOjORjLѭDQKL
KVӕQJӣQKLӋW
o
o
30 ±45 &QKѭQJFNJQJFyQKLӅXORjLѭDQ
C
[13, 18].
ĈDVӕ
Bacillus VLQKWUѭӣQJӣS+ PӝWV
±10
QKѭ
Bacillus alcalophillusKD\FyORҥLSK
KӧSYӟLS+
±QKѭ
Bacillus

acidocaldrius.
Bacillus Fy NKҧ QăQJ VҧQ VLQK QKLӅX H
SURWHDVH FHOOXODVH«
 GR ÿy FK~QJ
QJKLӋSWURQJEҧRYӋP{LWUѭӡQJ«0ӝ
ѭӡQJJһSWUR
QKLrQ
Bacillus subtilis, Bacillus megaterium, Bacillus mensentericus,
Bacillus pumilus«>
].

+uQK

1.1. +uQK̫QKQKX͡P*UDPP͡WV͙
4


7әQJTXDQYӅ[DNKXҭQ
Streptomyces
;ҥNKXҭQOjQKyPOӟQYLNKXҭQJUDP
ӃXNKtVӕQJKR
Fy FҩX WҥR GҥQJ VӧL SKkQ QKiQK ;ҥ
WUѭӡQJÿҩWWKDP JLD YjRTXi WUuQK
WURQJÿҩW
;ҥ NKXҭQ FQJ QKyP YӟL YL
rokaryota)
NKXҭQ F
WKXӝF
JLӟLNKӣLVLQK
Monera).
;ҥNKXҭQFyQKLӅXÿһFÿLӇPJLӕQJY
- .tFKWKѭӟFFӫD[ҥNKXҭQQKӓEpWѭѫ
- 1KkQFӫD[ҥNKXҭQOjQKkQQJX\rQWK
FKѭDFyGҥQJQKkQS
WKiL
- 7KjQKWӃEjRFӫD[ҥN
KXҭQNK{QJFKӭDFHOOXORVH
glucopeptit.
- 6ӵSKkQFKLDWӃEjRFӫD[ҥNKXҭQW
WKҩ\ӣYLNKXҭQ
- ;ҥNKXҭQNK{QJFyJLӟLWtQKÿ
ӵFFiLKRһFFKѭDU}JL
].

;ҥNKXҭQFyWKӇVLQKWәQJKӧSQKLӅ
PWUDRÿәLFKҩW
ÿһFELӋWFyêQJKƭDOjFiFFKҩWNKiQ
ELӃWFyÿӃQFyQJXӗQJӕFWӯ[ҥNKXҭQ
WKXӝF QKyP WHWUDF\FOLQHV PDFUROLGHV
NKҧ
QăQJVLQKUDFiFHQ]\PHPӝWVӕYL
+DL QKyP [ҥ NKXҭQ TXDQ WUӑQJ Oj W
Mycobacterium tuberculosis Jk\ EӋQK
Corynebacterium
ODR Yj
diphtheriae
Jk\ EӋQK EҥFK KҫX 0ӝW Vӕ HULD
[ҥ NKXҭ
Yj
&RU\QHEDFWHULDVӕQJFӝQJVLQKӣÿӝQJ
7UѭӟF ÿk\ [ҥ NKXҭQÿѭӧF [ӃS FKXQJ
WKӭFSKiWWULӇQGҥQJVӧLSKkQQKiQK
NKXҭQWKұWGRFK~QJFyQKLӅXÿһFÿL
YӟLQҩP
QKѭVDX
- &yJLDLÿRҥQÿDEjRYjÿѫQEjR
- .tFKWKѭӟFUҩWQKӓ
- 7KӇQKkQOjQKkQQJX\rQWKӫ\
- 9iFKWӃEjRNK{QJFKӭDFHOOXOR]HK
- .K{QJFyJLӟLWtQK
- 6ӕQJKRҥLVLQKKRһFNêVLQK>
].
;ҥNKXҭQSKkQEӕUӝQJUmLWURQJW
QѭӟFUiF

WKұPFKtFyWURQJFҧQKӳQJFѫFKҩWP
ÿѭӧF;ҥNKXҭQOjPӝWWURQJQKӳQJOR
QKLrQWtFKFӵFWKDPJLDYjRFiFTX
5


ÿҩWQѭӟFÿһF
ELӋWTXDQWUӑQJQKҩWFӫD[ҥNK
NKiQJVLQK
- [ҥNKXҭQSKkQOұSWӯÿҩW
WURQJVӕKѫQFKҩWNKiQJVLQKÿ
[ҥ NKXҭQ WURQJ ÿy 
ҥ Fy
NKXҭQ
QJXӗQ
KLӃP
J
Micromonospora YY« &iF [ҥNKXҭQKLӃP FXQJ
FyJLiWUӏFKLӃPWӟLFiFFKҩWNK
KӑFQKѭJHQWDPL[LQWREUDPL[LQYY«
1JRjLNKҧQăQJKuQKWKjQKFKҩWNK
FyNKҧQă
VҧQVLQKKjQJORҥWFiFFKҩWKӳXFѫ
FyJLiWUӏQKѭ9LWDPL
vitamin
%YLWDPLQ%YY«
FiFDFLGKӳXF
WUӏDVSDUDJLQLF
QKѭ
DODQLQH PHWLRQLQ Y

DP\ODVHD«
[3].
1.7әQJTXDQYӅQҩP
PөFWUҳQJ
Phanerochaete chrysosporium
Phanerochaete chrysosporium Oj PӝW WURQJ QKӳQJ Q
WUҳQJ WUrQ Jӛ Yj SKkQ Kӫ\ Jӛ ;XҩW
PҥQJQKӋQ+ӋWKӕQJHQ]\PHSKkQKӫ\
PFKRQyWU
ÿһFELӋW7KjQKSKҫQFӫDJӛFKӫ\ӃX
PjX QkX
Phanerochaete
/RjL
chrysosporium Fy PӝW Vӕ ÿһF ÿL
GөQJ
0
1y WKtFK QJKL YӟL(NKRҧQJ
QKLӋW
C) Fy
ÿӝ
 WKӇ
WѭѫQJ
Vӕ
WURQJQKӳQJÿӕQJӫF
yQKLӋWÿӝNKiFDR
1ҩP PөF WUҳQJWKѭӡQJ WuP WKҩ\ QKL
WUҫQ WURQJWӵQK
LrQ7KѭӡQJ WKu FiF ÿѫQYӏV\
F{QJKѫQFiFÿѫQYӏJXDLDF\O*
7

QKLӅXOLJQLQ*
WURQJNKLӣYiFKWӃEjRWKӭ
.K{QJJLӕQJQKѭFHOOXORVHYjKHPL
WKҷQJ KjQJ Pj Oj PӝW SRO\PHU WKѫP N
QKӳQJÿѫQYӏSKHQ\O
SURSDQRO1ҩPPөFWUҳQJÿѭӧ
QKѭOjQKӳQJ
YұWFyWKӇSKkQKӫ\KRjQWRjQOLJQLQ
ÿk\QJѭӡLWDTXDQWkPQKLӅXÿӃQVLQK
SKkQ Kӫ\ OLJQLQ WK{QJ TXD QKӳQJ QJK
Phanerochaete chrysosporium.
+Ӌ WKӕQJ SKkQ
Phanerochaete
Kӫ\ OLJQLQ
chrysosporium NK{QJ
FӫD
Eӏ
FҧPӭQJEӣLOLJQLQPjQy[XҩWKLӋQ
TXiWUuQKQX{LFҩ\WӭFOjVӵWăQJW
WKjQKSKҫQGLQKGѭӥQJ
6ӵFKX\ӇQKyDWKӭFҩSNKӣ
LÿҫXNKLFyV
FӫDFKҩW
FiF
QKѭ QLWURJHQ FDUERQ KD\
SKRVSKRU6ӵÿLӅXKzDFKX\ӇQKyDWKӭ
OLrQNӃWYӟLFKX\ӇQKyDJOXWDPDWH
LFҩ\QҩPWURQJP{
JJLӟL
6



KҥQQLWURJHQWKuVӵSKkQKӫ\OLJQLQ
[ҧ\UD
7URQJVӵNLӋQQj\
(
mQJKLrQFXUQJ
VSKkQK\OLJQLQOjPW
FSGRKjPOQJQLWURJHQFDJWK
WQF{QJJQLWURJHQV
EWKLXYjVFKX\QKy
K\OLJQLQVEWX
1.2. 71* 48$1 ơ1+
9 1*
10 9ơ 48< 75ẻ1+
751*
10758<17+1*
1.2.1. 7QJTXDQYQJ
jQKQPQFWD
1F WDFyWLPQQJOQYVQ[
ngXQQJX\rQOLXWUQJQPSKRQJSK~
LX NLӋQ WKӡL WLӃW WKXұQ OӧL FKR S
WUӗQJ QҩP TXDQK QăP &K~QJ WD ÿm Fѫ
JLӕQJYjVҧQ[XҩWQҩPÿӕLYӟLFiFO
PFKӫOӵFWKӏWUѭ
QJj\FjQJUӝQJPӣ&KtQKYuYұ\QJj
EDQKjQK 4X\ӃW
- 77J
ÿӏQK
ÿѭD

4Ĉ
QҩP ăQ QҩP
PөFVҧQSKҭPTXӕFJLDÿѭӧFѭXWLrQÿ
7ӱ
&KtQK3Kӫ1ѭӟF&ӝQJ+zD;m+ӝL&Kӫ
7KӡLJLDQTXDÿmFyQKLӅXP{KuQK
ӣTX\P{KӝJLDÿuQKWUDQJWUҥLJL
VҧQ [XҩW Vѫ FKӃ FKӃW
ELӃQ
QҩP ÿDQJ
NLQKWӯ
GR
SKiWWULӇQWKHRKѭӟQJFKX\rQQJKLӋS
NKkXVҧQ[XҩWVѫFKӃEҧRTXҧQWL
YӳQJ ÿҥW KLӋX TXҧ NLQK WӃ FDR WҥR
GkQĈӗQJWK
ӡLYLӋFSKiWWULӇQQJjQKQҩP
QkQJFDRJLiWUӏJLDWăQJFӫDVҧQ[
FӫDWUӗQJWUӑW
1.2.2. 7uQKKuQKVҧQ[XҩWYjWLrXWKө
1Jj\QD\ÿmSKiWKLӋQWUrQ
WURQJÿyFyNKR
Fy WKӇ ăQ ÿѭӧF Yj QX{L WUӗQJ WKjQK
UѫPQҩPPӝFQKƭQҩPNLPFKkPQҩP
YӵFGѭӧFOLӋXQKѭQҩPOLQKFKLQҩP
&yWUrQTXӕ
FJLDYQJOmQKWKәWUӗQJQҩ
NKRҧQJWULӋXWҩQQăP
WӕFÿӝWăQJWUѭӣQJEuQKT
- QăP&

QѭӟF VҧQ [XҩW QҩP KjQJ ÿҫX WKӃ JLӟL
WҩQWURQJÿyĈjL/RDQ
WәQJVҧQ
QҩPWKӃJLӟL+RD.ǤWҩQ
3KiS  WҩQ ,QGRQHVLD 
WҩQéWҩQ&DQDGD
2012)
7